:root{color-scheme:dark;--bg:#091e25;--surface:#112b33;--surface-2:#18363e;--text:#f2f6f5;--muted:#b6c8cc;--line:#2e4850;--lime:#d5ff9b;--violet:#b398ff;--width:1180px;font-family:Inter,"Segoe UI",Arial,sans-serif;font-size:16px}
*{box-sizing:border-box}html{scroll-behavior:smooth;scroll-padding-top:110px}body{margin:0;background:var(--bg);color:var(--text);line-height:1.65}a{color:inherit;text-decoration:none}a:hover{color:var(--lime)}button,input,summary{font:inherit}button,a,summary,input{-webkit-tap-highlight-color:transparent}a:focus-visible,button:focus-visible,summary:focus-visible,input:focus-visible{outline:3px solid var(--lime);outline-offset:5px}img{max-width:100%;display:block}h1,h2,h3,p{margin-top:0}h1,h2,h3{font-weight:500;line-height:1.14;letter-spacing:-.045em}h1{font-size:clamp(2.8rem,5.8vw,5.4rem)}h2{font-size:clamp(2rem,3.3vw,3.2rem)}h3{font-size:1.5rem}p{color:var(--muted)}.wrap{width:min(var(--width),calc(100% - 64px));margin-inline:auto}.skip{position:fixed;top:-100px;left:20px;z-index:100;background:var(--lime);color:var(--bg);padding:12px}.skip:focus{top:10px}.site-header{border-bottom:1px solid var(--line);background:var(--bg);position:relative;z-index:20}.nav-row{min-height:92px;display:flex;align-items:center;justify-content:space-between;gap:24px}.wordmark{font-size:1.7rem;font-weight:550;letter-spacing:-.06em;white-space:nowrap}.wordmark span{font-weight:350;color:var(--lime)}nav{display:flex;gap:30px;align-items:center;font-size:.94rem}nav>a{padding-block:20px}nav>a[aria-current=page]{color:var(--lime)}.nav-group{position:relative}.nav-group summary{cursor:pointer;list-style:none;padding:20px 0}.nav-group summary::after{content:'⌄';margin-left:8px;color:var(--muted)}summary::-webkit-details-marker{display:none}.nav-pop{position:absolute;top:58px;left:-18px;min-width:215px;background:var(--surface-2);border:1px solid var(--line);border-radius:14px;padding:10px;box-shadow:0 16px 40px #0005}.nav-pop a{display:block;padding:9px 12px;border-radius:6px}.nav-pop a:hover{background:var(--bg)}.contact-nav{border:1px solid #50707b;border-radius:40px;padding:10px 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0;list-style:none}.accordion summary::after{content:'+';font-size:1.8rem;font-weight:300;color:var(--lime);align-self:center}.accordion[open] summary::after{content:'−'}.accordion strong{display:block;font-size:1.1rem;font-weight:500}.accordion .meta{display:block;color:var(--muted);font-size:.88rem;margin-top:4px}.accordion-body{padding:0 42px 24px 0;color:var(--muted);font-size:.96rem}.accordion-body p:last-child{margin-bottom:0}.accordion-body li{margin-bottom:8px}.research-layout{display:grid;grid-template-columns:1.25fr 1fr;gap:70px}.list-heading{font-size:1.35rem;letter-spacing:-.02em}.search{display:block;width:100%;border:1px solid #4e6972;border-radius:12px;background:var(--surface);color:var(--text);padding:15px 18px;margin:12px 0 12px}.results-label{font-size:.82rem;color:var(--muted);margin:12px 0 24px}.paper{border-top:1px solid var(--line);padding:22px 0}.paper .journal{font-size:.8rem;letter-spacing:.02em;color:var(--violet);margin:0 0 7px}.paper h3{font-size:1.12rem;letter-spacing:-.015em;line-height:1.5;margin-bottom:10px}.paper-link{color:var(--lime);font-size:.85rem}.citation{font-size:.78rem;color:var(--muted);margin-top:12px}.citation summary{cursor:pointer}.citation p{margin:8px 0}.story-empty{border:1px dashed var(--line);border-radius:24px;min-height:290px;display:flex;flex-direction:column;justify-content:center;align-items:center;text-align:center;padding:40px}.story-empty p{max-width:280px;margin:16px 0 0;font-size:.94rem}.empty-mark{font-size:2rem;color:var(--violet);font-weight:300}.subnav{display:flex;gap:10px;flex-wrap:wrap;margin-top:26px}.subnav a{border:1px solid var(--line);border-radius:30px;padding:7px 16px;font-size:.88rem}.subnav a[aria-current=page]{background:var(--lime);border-color:var(--lime);color:var(--bg)}.activity{padding:28px 0;border-top:1px solid var(--line);display:grid;grid-template-columns:110px 1fr;gap:32px}.activity time{color:var(--violet);font-size:.9rem}.activity h2{font-size:1.4rem;letter-spacing:-.025em;margin-bottom:10px}.activity p{margin-bottom:0}.site-footer{border-top:1px solid var(--line);padding:65px 0 24px;background:#07191f}.footer-grid{display:grid;grid-template-columns:.8fr 1.4fr 1fr;gap:40px}.footer-grid h2{font-size:1.6rem;letter-spacing:-.04em;margin-bottom:14px}.email{color:var(--lime);font-size:1rem;overflow-wrap:anywhere}.footer-label{color:var(--text);font-size:1rem;margin-bottom:16px}.socials{display:flex;flex-direction:column;gap:10px}.socials a{display:flex;gap:12px;align-items:center;font-size:.9rem;color:var(--muted)}.social-symbol{width:25px;height:25px;display:grid;place-items:center;border:1px solid #789098;border-radius:5px;font-size:.72rem;color:var(--text);font-weight:700}.social-symbol.orcid{border-radius:50%;font-weight:500}.footer-bottom{display:flex;justify-content:space-between;gap:20px;margin-top:60px;padding-top:20px;border-top:1px solid var(--line);font-size:.8rem;color:#8da6ad}.draft-note{background:#18313a;color:#c5d7d9;text-align:center;padding:8px 18px;font-size:.75rem;border-bottom:1px solid var(--line)}[hidden]{display:none!important}.screen-reader{position:absolute;width:1px;height:1px;overflow:hidden;clip-path:inset(50%)}
@media(max-width:900px){.hero-grid{gap:35px;grid-template-columns:1.2fr 1fr}.hero{padding-block:65px}.wrap{width:calc(100% - 40px)}nav{gap:18px}.research-grid{gap:18px}.research-card h2{font-size:1.35rem}.research-layout{gap:32px}.about-section{gap:30px;grid-template-columns:.8fr 1.5fr}.footer-grid{grid-template-columns:1fr 1fr}.footer-grid>.wordmark{grid-column:1/-1}}
@media(max-width:650px){.nav-row{min-height:76px;flex-wrap:wrap;gap:0}.menu-toggle{display:block;color:var(--text);background:none;border:1px solid var(--line);border-radius:8px;padding:7px 12px}.js nav{display:none}.js nav.open{display:flex}nav{width:100%;padding:12px 0 24px;align-items:stretch;flex-direction:column;gap:0}nav>a,.nav-group summary{padding:11px 0}.nav-pop{position:static;box-shadow:none;margin-bottom:8px}.contact-nav{text-align:center;margin-top:10px}.hero-grid,.research-grid,.research-layout,.about-section,.footer-grid{grid-template-columns:1fr}.hero{padding:48px 0}.hero h1{font-size:3.2rem}.positions{grid-template-columns:minmax(0,1fr);gap:16px}.position{padding:19px 16px;border-radius:15px}.position h2{font-size:1.2rem}.position p,.position small{font-size:.78rem}.hero-intro{font-size:1rem}.research-card p{min-height:0}.research-card{margin-bottom:22px}.section{padding-block:54px}.page-head{padding-top:48px}.about-section{gap:12px;margin-bottom:48px}.research-layout{gap:45px}.footer-grid{gap:32px}.footer-bottom{flex-direction:column;margin-top:38px;gap:8px}.activity{grid-template-columns:1fr;gap:8px}.image-slot{max-height:320px}.research-card h2{font-size:1.65rem}}
.hero::before{background-image:url('images/banner.webp');background-position:center bottom;opacity:.65}
.research-figure{background:#fff}
.lab-name{color:var(--lime);font-size:1.05rem;line-height:1.55;max-width:580px;margin-bottom:16px;letter-spacing:.01em}
.home-service{padding:52px 0 100px;position:relative;isolation:isolate}
.home-service::before{content:"";position:absolute;inset:0 -16px;z-index:-1;pointer-events:none;background:radial-gradient(ellipse at 50% 58%,#7855bb30 0%,#73869315 36%,transparent 68%)}
.home-service .about-section{display:block;max-width:850px;margin:0 auto}
.home-service .about-section>h2{text-align:center;font-size:clamp(1.8rem,3vw,2.15rem);margin:0 0 48px;letter-spacing:-.035em}
.home-service .accordion,.home-service .accordion:first-child{background:#05051d;border:1px solid #ffffff08;border-radius:20px;margin-bottom:12px}
.home-service .accordion summary{padding:23px 32px;gap:24px;align-items:center;border-radius:20px}
.home-service .accordion summary strong{font-size:clamp(1.05rem,1.8vw,1.4rem);line-height:1.4}
.home-service .accordion summary::after{display:grid;place-items:center;width:40px;height:40px;border-radius:50%;background:#17172e;color:#e2dcf2;font-size:1.65rem;line-height:1}
.home-service .accordion summary:hover{background:#ffffff04}
.home-service .accordion .meta{font-size:.9rem;margin-top:6px;color:#aab7c3}
.home-service .accordion-body{padding:0 96px 26px 32px;font-size:1rem}
@media(max-width:650px){.home-service{padding:32px 0 64px}.home-service .about-section>h2{margin-bottom:30px}.home-service .accordion summary{padding:20px;gap:14px}.home-service .accordion summary::after{width:34px;height:34px}.home-service .accordion-body{padding:0 20px 24px}}
.accordion-body strong{color:var(--text);font-weight:600}
.research-figure img{object-fit:contain;padding:10px}
.research-figure[href="one-health.html"] img{position:absolute;top:50%;left:0;width:100%;height:125%;padding:0;transform:translateY(-50%);object-fit:contain}
.research-figure .card-arrow{bottom:8px;right:8px;width:27px;height:27px;box-shadow:0 1px 5px #0002}
.featured-talk{display:grid;grid-template-columns:minmax(0,.8fr) minmax(0,1.2fr);gap:clamp(2rem,5vw,5rem);align-items:center;border-top:1px solid var(--line)}
.featured-talk-copy h2{font-size:clamp(1.8rem,3vw,3rem);max-width:16ch;margin:.5rem 0 1rem}
.featured-talk-copy>p:not(.eyebrow){color:var(--muted);max-width:52ch}
.video-frame{position:relative;aspect-ratio:16/9;overflow:hidden;border:1px solid var(--line);border-radius:24px;background:#07151b;box-shadow:0 18px 48px #0004}
.video-frame iframe{position:absolute;inset:0;width:100%;height:100%;border:0}
@media(max-width:650px){.featured-talk{grid-template-columns:1fr}.featured-talk-copy h2{max-width:none}.video-frame{border-radius:18px}}
@media(prefers-reduced-motion:reduce){html{scroll-behavior:auto}}
.art-gallery{display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:3rem}.art-card img,.art-card video{display:block;width:100%;max-height:36rem;object-fit:contain;background:transparent}.art-card h2{font-size:1.6rem}.art-card p{max-width:65ch}.art-card .meta{margin-top:1.5rem}@media(max-width:700px){.art-gallery{grid-template-columns:1fr}}
.story-list{display:grid;gap:24px}.press-card{border:1px solid var(--line);border-radius:22px;background:var(--surface);overflow:hidden}.press-card>a{display:block;padding:30px}.press-card:hover{border-color:var(--violet)}.press-card h3{font-size:1.55rem;line-height:1.3;letter-spacing:-.03em;color:var(--text);margin-bottom:18px}.press-card .press-source{font-size:.875rem;color:var(--violet);margin-bottom:18px}.press-card .press-summary{font-size:1rem;line-height:1.7;margin-bottom:24px}.press-card .paper-link{font-size:.95rem}@media(max-width:650px){.press-card>a{padding:24px}}
.research-stories{display:grid;gap:32px}.research-story h3{font-size:1.3rem;line-height:1.4;letter-spacing:-.025em;margin:0 0 14px}.story-image{display:block;position:relative;aspect-ratio:1.6;overflow:hidden;border-radius:20px;background:var(--surface);border:1px solid var(--line)}.story-image img{width:100%;height:100%;object-fit:cover}.story-image:hover{border-color:var(--lime)}.research-story .story-credit{font-size:.8rem;line-height:1.5;margin:9px 0 0;color:var(--muted)}
/* Homepage typography follows the compact Poppins treatment of mol.org. */
@font-face{font-family:Poppins;src:url('fonts/poppins-medium.ttf') format('truetype');font-style:normal;font-weight:500;font-display:swap}
.hero h1{font-family:Poppins,"Segoe UI",sans-serif;font-size:42px;font-weight:500;line-height:1.1;letter-spacing:-.04em;margin-bottom:20px}
.hero .name-cn{font-family:"Segoe UI",sans-serif;font-size:24px;line-height:1.25;letter-spacing:-.02em;margin-top:12px}
.hero .lab-name{font-family:Poppins,"Segoe UI",sans-serif;font-size:18px;font-weight:500;line-height:1.45;letter-spacing:-.02em;max-width:480px;margin-bottom:32px;color:var(--text)}
.hero .position small,.hero .position p{font-size:14px;line-height:1.4}
@media(min-width:1440px){.hero h1{font-size:56px}}
@media(max-width:650px){.hero h1{font-size:38px}.hero .lab-name{font-size:16px}.hero .name-cn{font-size:22px}.hero .position small,.hero .position p{font-size:13px}}
@keyframes hero-arrive{from{opacity:0;transform:translateY(22px)}to{opacity:1;transform:translateY(0)}}
@keyframes hero-background{from{transform:scale(1.055);opacity:.35}to{transform:scale(1);opacity:.65}}
@media(prefers-reduced-motion:no-preference){
.hero::before{animation:hero-background 1.6s cubic-bezier(.2,.65,.3,1) both}
.hero h1,.hero .lab-name,.hero .pill,.hero .position{animation:hero-arrive .85s cubic-bezier(.2,.65,.3,1) both}
.hero h1{animation-delay:.1s}.hero .lab-name{animation-delay:.22s}.hero .pill{animation-delay:.34s}
.hero .position:first-child{animation-delay:.25s}.hero .position:last-child{animation-delay:.4s}
}
/* Reference desktop type scale; retain readable intermediate/mobile sizes. */
@font-face{font-family:InterNav;src:url('fonts/inter-medium.woff2') format('woff2');font-weight:500;font-style:normal;font-display:swap}
@font-face{font-family:Satoshi;src:url('fonts/satoshi-medium.woff2') format('woff2');font-weight:500;font-style:normal;font-display:swap}
.site-header nav{font-family:InterNav,Inter,sans-serif;font-size:18px;font-weight:500;line-height:24px;letter-spacing:.15px}
.site-header .wordmark{font-family:"Segoe UI Light","Segoe UI",Arial,sans-serif;font-size:clamp(1.5rem,2.3vw,2rem);font-weight:300;line-height:1;letter-spacing:-.045em;color:var(--text)}
.contact-nav,.hero .pill{font-size:16px;line-height:1.8;letter-spacing:-.2px}
.hero h1{font-size:clamp(2.625rem,4.75vw,5.5rem);line-height:1.1;letter-spacing:-.04em}
.hero-grid>div:first-child{container-type:inline-size}
.hero .lab-name{font-family:Satoshi,Arial,sans-serif;font-size:clamp(1rem,5.4cqi,3.5rem);line-height:1.3;letter-spacing:-.02em;max-width:none}
.hero .lab-name span{display:block}
.hero .wrap{width:min(1600px,calc(100% - clamp(40px,6vw,128px)))}
.hero-grid{grid-template-columns:minmax(0,1.5fr) minmax(0,1fr);gap:clamp(32px,5vw,90px)}
.hero-grid>div{min-width:0}
.hero .position small,.hero .position p{font-size:16px;line-height:1.6;letter-spacing:-.02em}
.page-head p:not(.eyebrow){font-family:Satoshi,Arial,sans-serif;font-size:27px;line-height:1.4;letter-spacing:-.02em}
.research-card p{font-family:Satoshi,Arial,sans-serif;font-size:16px;line-height:1.4;letter-spacing:-.02em}
@media(max-width:1100px){.page-head p:not(.eyebrow){font-size:24px}.site-header nav{gap:18px}}
@media(max-width:760px) and (min-width:651px){.nav-row{gap:12px}.site-header nav{gap:12px;font-size:16px}.contact-nav{padding:8px 14px}}
@media(max-width:650px){.hero-grid{grid-template-columns:1fr}.page-head p:not(.eyebrow){font-size:20px}.hero .position small,.hero .position p{font-size:14px}.site-header nav{font-size:18px}.menu-toggle{font-size:18px}}

/* Research figures use the same thumbnail frame and overlaid arrow as other stories. */
.story-image[data-fit=contain]{aspect-ratio:1.6;border-radius:20px}.story-image[data-fit=contain] img{height:100%;object-fit:cover}

.artwork-index{padding-block:48px 80px;display:grid;grid-template-columns:repeat(2,minmax(0,1fr));gap:48px}.artwork-cover{display:block;width:100%;max-width:480px;justify-self:center}.artwork-cover img{width:100%;height:auto;transition:transform .25s ease}.artwork-cover:hover img{transform:translateY(-4px)}.artwork-detail{padding-block:42px 90px}.artwork-back{display:inline-block;margin-bottom:32px}.artwork-title h1{font-size:clamp(2.5rem,5vw,4.6rem);margin-bottom:44px}.artwork-layout{display:grid;grid-template-columns:1.15fr 1fr;gap:56px;align-items:start}.artwork-layout figure{margin:0}.artwork-layout img{width:100%;height:auto}.artwork-copy p{font-size:1rem;line-height:1.9}.artwork-copy section{border-top:1px solid var(--line);margin-top:30px;padding-top:28px}.artwork-copy h2{font-size:1.5rem;letter-spacing:.04em;margin-bottom:18px}.artwork-copy [lang=zh] p{line-height:2;letter-spacing:.025em}@media(max-width:800px){.artwork-layout{grid-template-columns:1fr;gap:32px}.artwork-copy{max-width:65ch}.artwork-index{grid-template-columns:1fr;padding-block:28px 56px}.artwork-title h1{margin-bottom:28px}}@media(prefers-reduced-motion:reduce){.artwork-cover img{transition:none}.artwork-cover:hover img{transform:none}}

.latest-news{display:flex;align-items:center;gap:22px;padding:18px 0;border-bottom:1px solid var(--line);font-size:.9rem}.latest-label{color:var(--lime);white-space:nowrap}.latest-news time{color:var(--muted);font-size:.85rem;white-space:nowrap}.latest-title{flex:1}.latest-news:hover .latest-title{text-decoration:underline;text-underline-offset:4px}@media(max-width:650px){.latest-news{display:grid;grid-template-columns:1fr auto;gap:8px 16px}.latest-title{grid-column:1}.latest-news>span:last-child{grid-column:2;grid-row:2}}